Field of Science

Showing posts with label taxonomy. Show all posts
Showing posts with label taxonomy. Show all posts

You still say tomato

Tomato, tomato (said with long and short vowel sounds), a domesticated solanaceous fruit that by any other name would still taste as good, especially while thinking about the sugo alla puttanesca the Phactor cooked last night from fresh Amish paste tomatoes. For quite some time, botanists called the tomato, Lycopersicon esculentum, the juicy wolf peach, a name which added a bit of European skepticism about how esculent this neotropical, and newly arrived, nightshade might be. If you know anything about Old World nightshades, then you'll understand the skepticism. No one doubted that the tomato was closely related to the huge genus Solanum, the name sake of the nightshade family, and now relatively recent molecular studies have shown that the tomato species is part of the genus Solanum. Now what is usually done in these circumstances is that the specific epithet (esculentum) is transferred back to Solanum to produce a new combination, Solanum esculentum. Ta da! And for awhile that was the species name of tomato, but then someone remembered Linnaeus. Remember Linnaeus? The father of taxonomy. Well, the order in which taxonomic names are published counts with the first name published (the oldest) being judged correct, and no one is older than Linnaeus. Linnaeus was simply set as the starting date of plant names, and Linnaeus had named this plant Solanum lycopersicon. Subsequent to Linnaeus the specific epithet was raised to a generic level and a new specific epithet was added, but now that tomato is back in Solanum the whole thing reverts to Linnaeus' original species name. Image credit - diversely colored wild tomatoes from western S. America: Ana Caicedo, Univ. Mass.

Friday Fabulous Flower - but just for today!

As mentioned a few days back, members of the lily family, or members belonging to the diverse lineages that used to constitute the lily family, contribute a great deal to the color in our summer gardens. So here's a rather spectacular daylily from Mrs. Phactor's collection; the flower is easily 20 cm across. Unfortunately these gaudy giant flowers only last one day, thus the name.
Now the Phactor knows what you're thinking, "If a daylily is no longer in the lily family, what is it classified as?" The genus Hemerocallis in in the Xanthorrhoeaceae (zan-thor-ree-aye-see-ee), which isn't even in the Liliales but the Asparagales. So in addition to the Liliaceae, genera that all used to be dumped into this family or very closely related families like the Amaryllidaceae (Did you learn that the difference was an inferior ovary vs. a superior ovary?) there are now 13 more families here in alphabetical order, unless one got missed: Agavaceae, Alliaceae, Asparagaceae, Asphodelaceae, Hyacinthaceae, Melanthiaceae, Nartheciaceae, Ruscaceae, Smilacaceae , Tecophilaeaceae, Themidaceae, Tofieldiaceae, and Uvulariaceae. Wow! It used to be so much simpler and so much more incorrect! Some people ask, “Why bother, the old classification worked fine.” Well, that’s only true if all you needed was a classification that was nothing more than a card catalog for filing all these genera. However, modern plant taxonomy sees classifications as phylogenetic hypotheses, so groupings hypothesize a common ancestry for that particular lineage, and as is clear, although no so the relationships among all these lineages for which you need a diagram, the “lily lineage” represents a great deal of evolutionary diversity. If all this is just too much, the flower is just as pretty.

Time to update my taxonomy and YOURS

One of the strengths and problems with science is that it keeps finding out new things, and my job is to keep up. But at times it's a struggle because you have to relearn things you've KNOWN for more than 40 years. So let's say you get a book on trees and it's organized alphabetically by botanical families. No problem, mostly, to find a maple you open near the front looking for Aceraceae, but its not there. Maybe you want to find a buckeye; no problem, you've known the family name forever, but you can't find Hippocastanaceae anywhere. What kind of a book on trees is this anyway? Well, it's up to date so Acer and Aesculus are located in the Sapindaceae, the soapberry family. This may well come as a surprise but the research was published 5 years ago. Now in the old days such taxonomic rearrangements might have bordered on whimsy, and some might become popular and others forgotten, but now phylogenetic research clearly demonstrates that a broader family concept should apply and these two prominent northern temperate genera long residing in their own families should be with the other soapberries. Unfortunately for those of us who are the go-between for putting the scientific literature into the classrooms and blogs, it's nearly impossible to keep up with the changes especially in areas outside of your research expertise(s). The Phactor will try to adjust his thinking, but it's just like when meeting a former student, and impressively the free-wheeling memory access process pulls up the name - Goldschmidt, "Hey, Ms. Goldschmidt, how nice to see you." She's impressed her old professor got the name right, but she's married now to an Abercrombe. Well, no way in ever loving memory enhancers will she ever get reclassified. Constantly having to update all those connections becomes a real problem. But it can be worse; a colleague used to complain to me that she couldn't start and finish a project on any legume genera without having the name change.

Know your genera - Lesson 4: Chrysanthemum

More good news! You know yet another scientific name of a plant: Chrysanthemum. But, it isn't the plant you think it is. Those darned taxonomists figured out that chrysanthemums (chryos- Greek for gold; anthemon - Greek for flower) did not form a natural group and various species have been segregated into about 10 different genera.
The familiar "florists' chrysanthemums" pictured here and so popular this time of year are now in the genus Dendranthema (dendro = woody, a reference to their woody annual stems). Like many other people, the Phactor learned to call the Shasta or ox-eye daisy Chrysanthemum lecanthemum, which is kind of fun to say, except it meant "goldflower-whiteflower". Leucanthemum (leuco = white) is their new genus. Other former chrysanthemums are the pyrethrum daisies (feverfew - long known as a remedy for fevers, and as a source of the insecticide of the same name) are now placed in Tanacetum. And what of the real chrysanthemums? Of the 300-400 species that used to be in this genus, now there are two, both annuals of the Mediterranean region.

A rose, is a rose, so how does I knows?

Our woody plant horticulturalist stopped by yesterday with a piece of leafy twig. "What do you think this is?" he asked. Hmm, after assessing the specimen, I said, "Well, it's in the rose family." He already had guessed that, but beyond that it wasn't obvious. Now this particular specimen turned out to be a bit unusual, a species I had never seen before, and if that were not the case, he wouldn't have needed my input at all. This particular specimen proved to be a tree quite uncommon in this area (Sorbus aria, whitebeam). After visiting several references, several possibilities were eliminated, and it wasn't until I tried an old woody plant identification key, one that includes ornamental species, that things began to make sense. This species has a simple leaf and most species of Sorbus have pinnately compound leaves, a single axis with two rows of leaflets. Well and good, between the two of us, we nailed the ID and felt pretty good about ourselves.

But here's the thing. Neither one of us ever considered any other possibility after initially deciding this plant was a member of the rose family, instead of one of the other 700 or so families. Now the rose family is a pretty big group of plants, 100-120 genera and 3000 to 3500 species. What made us decide rose family?

It's strange but I don't actually know. Of course I can recite a list of characters, but most of them were not present because there were no flowers or fruit. Yet this twig, with its dozen and a half leaves and buds somehow just looked "rosy". My first thought was a pear, but this specimen's leaves had a double saw toothed margin (pear's are mostly smooth edged), wooly white hairs on the back side of the leaves (never seen a pear like that), and rounded buds (pears are usually more pointy). OK so not pear, more cherry like, but the bark was very un-Prunus, no horizontal lenticels and shiny buds. And finally by a process of elimination I ended up at Sorbus, even though the leaf seemed all wrong (simple leaf rather than pinnately compound). Once this hurdle was cleared, the details fell into place. Score one for the botanist.

This is the hard thing about plant identification. At a certain point, you have enough experience, that you can simply use a gestalt. Some sort of search image is triggered that shoves you in the right direction. And even though I have been teaching such courses for years, I just don't know how to teach this. You simply must work at identifying plants long enough and if you are good, this sense comes to you.

I play a dirty trick on students learning plant ID by giving them two very closely related plants sequentially, sometimes the same species, but just with different colored flowers or leaves. Some few look upon the 2nd specimen with a puzzled look, and then ask, didn't we just do this? Some are even more certain, and toss it aside as knowing I tried a lame trick. Others without an iota of recognition labor through another step by step slog through an identification key, and act surprised when the same species comes up again. Interestingly this exercise has proven to be a great predictor of their over all performance in the course. And I wonder if this skill, this perception, is tied to the ability to conceptualize, to go past the details and grasp the essential underlying idea. Because that's how this works with plant ID. The species ID is in the details, but the broader classification, in particular the family level taxonomic grouping is in the conceptualizing of the commonalities.

That's one good thing about experience. You do get better in doing some things with age. And you can't hold a specimen up to your computer monitor and get any closer to an ID. Technology isn't making any serious inroads into such skills at all. You can scan it, or digitally photograph it, and put it on the internet, but sooner or later, it's someone like me who tells you what it is.

A long time colleague of mine once expressed his concern and apparent inability to teach such skill. "Maybe we can just rip their heads off and pour it in," he said. Sounds like fun.