Field of Science

Showing posts with label horsetails. Show all posts
Showing posts with label horsetails. Show all posts

What? No strobili! That's OK, it goes with no observations

Plants can be so uncooperative.  The Equisetum (3 species) in our glasshouse was most untimely and no strobili were available (other than pickled) for lab.  Probably that was OK because my way too many of my students were in an uncooperative, non-observational mood.  They broke all the "make observations" rules.  Clicking pictures of things they didn't study is the worst, especially when the fossils are too small to see anything.  You might as well turn in an image of green water in a test tube and label it "Chlamydomonas"; it's been done, and it pretty well demonstrates that the student in question didn't observe a damned thing.  This is the down side of digital.  It promotes a superficial, fast and easy approach, a non-learning approach.  The don't pay attention attitude was so bad that at the end of class TPP had to re-sort all of the microscope slides because students didn't pay enough attention, read the labels, and match them with the correct slide trays.  Pure sloppiness.  Now of course this was not universal; the gap just widens between the attentive and the inattentive.  It makes it easy to sort the sheep from the goats, which is part of my job, but you wish you had enough time to promote better study habits.   

Horsetails are not ferns!

Common names cause a lot of grief, but they remain friendly, although often uninformative or misleading, easy to remember, and at times they are useful as shorthand references.  They certainly have their place in teaching and outreach.  Now here's the particular problem.  Molecular data nests the horsetails within the lineage of what traditionally have been called ferns.  OK, fair enough, but that does not make horsetails into ferns.  In a classification of extant (living) ferns published in 2006 my colleagues label the whole lineage "ferns".  Right now the Phactor does not wish to argue about whisk ferns (also called psilophytes) in this same context.  But horsetails have been  distinct from ferns back to the point when there were neither ferns nor horsetails, but merely a group of plants that were the likely common ancestors of ferns and horsetails, and even within this group their respective ancestors were different.  Traditionally, the lower vascular plants (those without seeds) were called the pteridophytes - the fern-like plants.  Now this is fine and everyone understood what was meant, but the beautiful people have decided that because pteridophytes used to include the clubmosses, the term must be avoided.  True, the clubmosses are a separate lineage, so you must re-define pteridophyte to exclude the clubmosses, and this type of thing is done all the time.  Pteridophyte always had a broader connotation than the term "fern"; it basically meant the "ferns and fern allies (which also included clubmosses)".  When you say fern, the image your mind forms is not at all what you see when someone says "horsetail".  Until people get used to the narrower definition of pteridophyte there will be some confusion, but certainly that produces no greater confusion than calling horsetails "ferns".  Part of the trouble is that the plants traditionally considered "ferns" do not form a single lineage; no lineage of ferns exists unless you include the whisk ferns and horsetails, but now you've got pteridophytes.  So the Phactor still sees considerable utility to pteridophyte to label the whole lineage, although now redefined to exclude clubmosses, because he cannot bring himself to call a horsetail a fern. If anyone wants this publication, go to this link (Kathleen Pryer's web page), scroll down to 2006; it's the 1st entry.  What think you RE ferns and horsetails?

Age of Equisetum

Presently the horsetail lineage, whose diversity peaked in the Carboniferous, has only a single genus, Equisetum. A newly described fossil (American Journal of Botany) indicates that the genus appeared much earlier than previously thought. The anatomical detail of these fossils is quite amazing (stem cross section) and clearly shows that this late Jurassic (ca. 150 million years ago) horsetail has the characteristics of Equisetum. While that's a really long time for a genus to be around, the champion is still the clubmoss Selaginella, a genus that appeared in the Carboniferous which would make it about twice as old as Equisetum. And that's rock hard evidence.